Hyphodontia griselinae

(G.H. Cunn.) E. Langer

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Synonyms:

= Rogersella griselinae (G.H. Cunn.) Stalpers 1985, N. Z. Journ. Bot. 23: 305.
= Corticium griselinae G. H. Cunn. 1963, N. Z. Dep. sci. industr. Res. Bull. 145: 71 (Basionym).
Holotype: New Zealand, Wellington, Mt. Tongariro, 900 m alt., on Griselina lucida, leg. G. H. Cunningham, Nr. 17424, Herbarium PDD.

= Rogersella asperula Liberta & Navas 1978, Can. J. Bot. 56: 1780.
Holotype: Venezuela, Parque Nacional El Avila, on rotten wood, No. 1640, Herbarium ISU.
Isotype: Herbarium ARIZ.

Macromorphology:

Resupinate, odontioid, chalky white, margin diffuse thinning out, aculei up to 50 µm high.

Micromorphology:

Spores:
4-5 µm in diameter, globose, verrucose. slightly thick-walled (< 0.5 µm), hyaline, inamyloid. TEM photo of spore ornamentation (60 K).
Basidia:
12-15 x 4-5 µm, suburniform with one basal clamp each.
Sterigmata:
Four, 4-5 µm long.
Cystidia:
Capitate cystidia, up to 30 µm long, 4-5 µm in diameter, basal ventricose, apikal with exsudat, which is soluble in 5% KOH solution.
Clamps:
All primary septa with one clamp each.
Hyphae:
Hyphal system monomitic. Subhymenial hyphae 2.5-3 µm in diameter, thin- walled , particularly totally encrusted with crystals. Basal hyphae 2-3 µm in diameter, slightly thick-walled, hyphae loosely running. Hyphal branching by subseptal outgrowingor by outgrowing of clamps.
Aculei:
Densely connected, capitate cystidia forming the aculei.

Habitate and substrate:

Dead parts of Dryopteris and Woodwardia.
Other substrats: Cryptomeria, Erica, Olearia, Griselina.

Biogeography:

All known specimens are from subtropical or tropical areas.

Remarks:

H. griselinae with verruccose spores in combination has typical features of the genus Hyphodontia (capitate cystidia, suburniform basidia, hyphodontioid hyphal branching). For this species Liberta & Navas (1978) described the genus Rogersella with the species R. asperula and and delimited it against Hypho-derma, Trechispora and Hypochnicium . Cunningham nevertheless described this species doubtless in 1963 as Corticium griselinae. This older name has to be used in priority.

H. griselinae is very close to H. sambuci and H. fimbriata . The fruit bodies of these three species are very strong incrusted with crystals. Because of this, the basidiocarp is like a chalky white surface on the substrate. From H. sambuci a aerophytical growth on branches or bushes is well known. The strong incrustion could be seen as a adaption to dry weather conditions. Unfortunately there aerophytical growth is not known of H. fimbriata and H. griselinae. Dolipores with continuous parenthesomes, hyphodontioid hyphae, and the characteristic droplet in the spores, let no doubt on the position of H. griselinae within the genus Hyphodontia.

The ornaments of the spores is a leading character of some taxa (Ganoderma: double spores wall with spinulose endospores; Thelephoraceae: Spores with double warts; Russula: amyloid spore ornamentic). On the other hand the genera Botryobasidium, Hyphodontia, Phlebiella, Trechispora or Tulasnella have as well species with smooth and ornamented spores within one genus. This appearently inconsistency in the importance of the feature "ornamented spores" depends on the different spectrum of characters of the respective taxa. Is it possible to detect an other leading character within a genus it is very well founded that species with ornamented and smooth spores are in the same genus like in Hyphodontia or Tulasnella.

Herbarium specimens:

Spain, Canary Islands, La Gomera, Cabezo Alto, 5 km sout east of Hermigua, in a decidous forest, on Woodwardia radicans, leg. L. Ryvarden, 16.01.1974, LR 12552, Herbarien ISU, O.
Spain, Canary Islands, Teneriffa, Pijaral Anaga, on Dryopteris sp., leg. L. Ryvarden, 18.02.1989, LR 26518, Herbarium O.
Spain, Canary Islands, Teneriffa, Pijaral Anaga, on Woodwardia sp., leg. L. Ryvarden, 18.02.1989, LR 26497, Herbarium O.
Spain, Canary Islands, Teneriffa, Relajos, Baranco de la Caldera, on Erica arborea, leg. F. Beltrain et al., 29.11.1987, TFC MIC 5919, Herbarium O.
La Réunion, Cilaos VII, under the crossing Roche merveilleuse - route de Bras Sec", 1370-1400 m alt., on Cryptomeria japonica, leg. J. Boidin, 14.04.1985, LY 11103, Herbarium LY.


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Dr. Ewald Langer - 22. Aug. 1995
gelanger@mailserv.zdv.uni-tuebingen.de